Physical Mechanism: Kelvin waves depress the thermocline from a baseline of ~25m down to 68.4m. Coastal upwelling winds can no longer dredge nutrient-rich cold sub-surface water. Engraulis ringens (anchoveta) schools plunge below avian plunging limits (10–15m), precipitating mass northern migration to the seasonal wind-jet upwelling in the Gulf of Panama.
While Humboldt upwelling collapses under positive Kelvin anomalies, the Panama Canal / Isthmus gap winds generate localized seasonal Ekman pumping in the Gulf of Panama (Jan–Apr), creating a fertile shallow pelagic feeding ground that shelters displaced Pacific seabird populations.